Woods Hole Oceanographic Institution
ATLAS - GAZETTEER COLLECTION

NOAA Technical Memorandum NMFS

^TES O* *

1988

ICHTHYOPLANKTON AND STATION DATA FOR

CALIFORNIA COOPERATIVE OCEANIC FISHERIES

INVESTIGATIONS SURVEY CRUISES IN 1975

David A. Ambrose
Richard L. Charter
H. Geoffrey Moser
Bradley S. Earhart

PLEAGE RETURN

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INSTITUTION DATA LIBRARY

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NOAA-TM-NMFS-SWFC-1 10

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U.S. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration

National Marine Fisheries Service

Southwest Fisheries Center

NOAA Technical Memorandum NMFS

The National Oceanic and Atmospheric Administration (NOAA), organized in
1970, has evolved into an agency which establishes national policies and
manages and conserves our oceanic, coastal, and atmospheric resources. An
organizational element within NOAA, the Office of Fisheries is responsible for
fisheries policy and the direction of the National Marine Fisheries Service
(NMFS).

In addition to its formal publications, the NMFS uses the NOAA Technical
Memorandum series to issue informal scientific and technical publications when
complete formal review and editorial processing are not appropriate or feasible.
Documents within this series, however, reflect sound professional work and may
be referenced in the formal scientific and technical literature.

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ATLAS ■ GAZETTEER COLLECTION

NOAA Technical Memorandum NMFS

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1988

ICHTHYOPLANKTON AND STATION DATA FOR
CALIFORNIA COOPERATIVE OCEANIC FISHERIES
INVESTIGATIONS SURVEY CRUISES IN 1975

David A. Ambrose

Richard L. Charter

H. Geoffrey Moser

Bradley S. Earhart

Southwest Fisheries Center

National Marine Fisheries Service

La Jolla, CA 92038

NOAA-TM-NMFS-SWFC-1 10

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U.S. DEPARTMENT OF COMMERCE
C. William Verity, Jr., Secretary

National Oceanic and Atmospheric Administration

William E. Evans, Under Secretary for Oceans and Atmosphere

National Marine Fisheries Service

James W. Brennan, Assistant Administrator for Fisheries

CONTENTS

Page

List of Figures iii

List of Tables iv

Abstract 1

Introduction 1

Sampling Area and Pattern 2

Sampling Gear and Methods 3

Laboratory Procedures 4

Identification 5

Computer Entry and Editing 9

Species Summary 10

Explanation of Tables 11

Acknowledgments 12

Literature Cited 14

Figures 18

Tables 29

Index 225

li

LIST OF FIGURES

Page

Figure 1. Composite arrangement of diagrammatic
charts showing areas sampled on each
CalCOFI cruise during 1975 18

Figure 2. Station pattern for CalCOFI Cruise 7412
showing track for the David Starr
Jordan 19

Figure 3. Station pattern for CalCOFI Cruise 7412

showing track for the Alexander Agassi z 2

Figure 4. Station pattern for CalCOFI Cruise 7501
showing track for the David Starr
Jordan 21

Figure 5. Station pattern for CalCOFI Cruise 7501

showing track for the Alexander Agassi z 22

Figure 6. Station pattern for CalCOFI Cruise 7503 23

Figure 7. Station pattern for CalCOFI Cruise 7505 24

Figure 8. Station pattern for CalCOFI Cruise 7507 25

Figure 9. Station pattern for CalCOFI Cruise 7510

and 7511 26

Figure 10. Station pattern for CalCOFI Cruise 7512 27

Figure 11. The basic station plan for CalCOFI

cruises from 1950 to the present 28

in

LIST OF TABLES

Page

Table 1. Station and plankton tow data for

CalCOFI cruises in 1975 29

Table 2A. Pooled occurrences of fish larvae taken
during the CalCOFI cruise in November-
December, 1974 67

Table 2B. Pooled occurrences of fish larvae taken

during CalCOFI cruises in 1975 69

Table 3A. Pooled numbers of fish larvae taken
during the CalCOFI cruise in November-
December, 1974 73

Table 3B. Pooled numbers of fish larvae taken

during CalCOFI cruises in 1975 75

Table 4. Numbers of fish larvae taken on stations

occupied during CalCOFI cruises in 1975 79

Table 5. Summary of pooled occurrences of fish
larvae taken on CalCOFI cruises from
1972-1981 219

Table 6. List of stations with multiple

occupancies in one month during 1975 224

IV

ABSTRACT

This report provides ichthyoplankton and associated station
and tow data from California Cooperative Oceanic Fisheries
Investigations (CalCOFI) cruises conducted off California and
Baja California in 1975. It is the twenty-first report in a
series that presents these data for all biological-oceanographic
CalCOFI surveys from 1951 to the present. A total of 1592
stations was occupied during eight monthly multivessel cruises
over the survey area which extended from Pt. Reyes, California to
Pt. San Juanico, Mexico, and seaward to several hundred miles.
The data are listed in a series of 6 tables; the background,
methodology, and information necessary for interpretation and
quantitative analysis of the data are presented in an
accompanying text. All pertinent station and tow data, including
volumes of water strained and standard haul factors, are listed
in the first table. Another key table lists, by station and
month, standardized counts of each of the 153 larval fish
categories identified from survey samples. This and previous and
subsequent reports make the CalCOFI ichthyoplankton and station
data available to all investigators and serve as guides to the
newly developed computer data base.

INTRODUCTION

This report, the twenty-first of a series, provides
ichthyoplankton and associated station and tow data from
California Cooperative Oceanic Fisheries Investigations (CalCOFI)
joint biological-oceanographic survey cruises conducted in 1975.
This program was initiated in 1949, under the sponsorship of the
Marine Research Committee of the State of California, to study
the population fluctuations of the Pacific sardine (Sardinops
sagax) and the environmental factors that may play a role in such
fluctuations. CalCOFI, known as the California Cooperative
Sardine Research Program from 1949 to 1953, was made up of
representatives of the South Pacific Fisheries Investigations
(SPFI) of the U.S. Fish and Wildlife Service [now the La Jolla
Laboratory, National Marine Fisheries Service (NMFS) ] , the
Scripps Institution of Oceanography (SIO) , the California
Department of Fish and Game (CDFG) , the California Academy of
Sciences (CAS) and the Hopkins Marine Station of Stanford
University. The first three of these agencies supplied ships and
personnel to conduct the sea surveys. NMFS processed the
plankton samples and analyzed the ichthyoplankton from them. SIO
processed and analyzed the hydrographic samples and measurements
and also analyzed invertebrate groups from the plankton samples.

The boundaries, station placement, and sampling frequency
for the CalCOFI survey area were based on the results of joint
biological and oceanographic cruises conducted by NMFS and SIO
during 1939-41. Those cruises were designed to collect sardine
eggs and larvae and associated hydrographic data over the entire

areal and seasonal spawning range of the species. On these
survey cruises, plankton tows were made to 70 m, a depth which
encompassed the vertical distribution of sardine eggs and larvae.
Wide-ranging joint biological and oceanographic survey cruises
were resumed in 1949 with sardine as the focus; however, an
increasing interest in other biological components resulted in
the deepening of standard tows to 140 m in 1951. This marked the
beginning of truly quantitative ichthyoplankton sampling on
CalCOFI surveys.

Some data resulting from CalCOFI surveys in 1975 have been
published. Hydrographic data (Univ. of Calif., SIO, 1984a, b)
were presented in standard formats. Distributional maps of
larvae of 2 taxa taken on CalCOFI surveys during 1975 are
presented in the CalCOFI atlas series: rockfish (Sebastes spp.),
Ahlstrom et al., 1978; and northern anchovy (Engraulis mordax) ,
Hewitt, 1980.

A computer data base for eggs and larvae of sardine and
anchovy, for larvae of Pacific hake (Merluccius productus) , jack
mackerel (Trachurus symmetricus) and Pacific mackerel (Scomber
japonicus), and for eggs of Pacific saury (Cololabis saira) was
established in 1969. The development of a data base for other
fish larvae is a complex undertaking because competency of
identification has evolved steadily over the past 38 years. We
began the task of producing a CalCOFI ichthyoplankton data base
and associated data report series in 1983. All available
original records for 1975 were subjected to an extensive
verification and editing process to produce this report. This
and previous (Ambrose et al., 1987a, b,c; 1988a,b; Sandknop et
al., 1987a,b; 1988a, b,c; Stevens et al . , 1987a, b,c; 1988a, b;
Sumida et al . , 1987a, b; 1988a, b,c) and subsequent reports make
the CalCOFI ichthyoplankton and station data available to all
investigators and serve as guides to the computer data base. The
data base will be modified when additional errors are discovered
and when composite taxa from the earlier years are reidentif ied.
These reports are the fundamental reference documents against
which subsequent changes in the data base can be compared.

SAMPLING AREA AND PATTERN

In 197 5, the seven CalCOFI survey cruises occupied stations
during portions of all months except April, August, and
September. An eighth cruise (7412) , conducted in November and
December of 1974, was included in the 1975 data base. A total of
1592 stations was included in this data base, with an average of
199 stations per cruise (range 50-279) . Coverage of the survey
station pattern varied among cruises and the entire survey area
was not covered on any single cruise (Figures 1-10, Table 1) .
The area off northern California (lines 40-57) was not covered.
All major lines were occupied off central California (lines 60-
77) on 7412, 7501, 7503, 7507, and 7511; only three lines were
occupied in this region on 7505. The area between Pt.
Conception, California, and Pt . San Juanico, Baja California

(lines 80-137) was occupied on 7412, 7501, 7507, 7510 and 7511;
southerly coverage of this region stopped at Sebastian Vizcaino
Bay (line 120) on 7505 and at Pt. Abreojos (line 130) on 7503.
The area off southern Baja California (lines 140-157) was not
surveyed in 1975. Typically, coverage did not extend beyond
station 90 (approximately 160-260 miles offshore) . Cruises 7510
and 7511 in our data base are considered as 7510 in the SIO
hydrographic data base and are combined in Figure 9.

Two vessels were employed on these cruises: the David Starr
Jordan of NMFS , and the Alexander Agassi z of SIO. The David
Starr Jordan was used on seven cruises and the Alexander Agassi z
on four (Table 1; Univ. of Calif., SIO, 1984a, b).

After 1969, CalCOFI surveys were made on a triennial basis.
These began in 1972 and continued every 3 years (1975, 1978,
1981, 1984) until 1985 when annual surveys were resumed.

SAMPLING GEAR AND METHODS

During 1975, a 1-m diameter ring net was used on all
cruises; the net was similar to that used on previous surveys
except the fabric was 0.505 mm nylon mesh instead of silk bolting
cloth (Smith, 1974). The cod end was constructed of 0.333 mm
nylon mesh. The frame was fastened to a short 3-lead bridle
connected to several meters of line which attached to the towing
cable by a clamp. A current meter was suspended in the center
of the mouth of each net to measure volume of water filtered
(see Kramer et al., 1972, for further details).

The standard tow in 1975 was an obligue haul to ca . 210 m
depth (to 15 m of the bottom in shallow areas) designed to filter
a constant amount of water per depth interval (ca. 3m /m of
depth) over the vertical range of most ichthyoplankters . Hauls
were made at a ship speed of 1.5-2.0 knots and initiated by

1
CalCOFI lines (Figure 11) are arranged perpendicular to the

coastline and extend from the Canadian border (line 10) to below

Cape San Lucas, Baja California (line 157). Stations were

established on the basis of a perpendicular to line 80 (off Pt.

Conception) at a point designated as station 60. Stations were

plotted seaward and shoreward from station 60 on each line.

Cardinal CalCOFI lines (those ending in "0") are 120 miles apart

and usually bracket two ordinal lines (ending in "3" or "7"), so

that lines are 40 miles apart over most of the pattern. Cardinal

stations are 40 miles apart and typically these are separated by

a station number ending in "5" so that stations are 20 miles

apart out to station 90 on most lines. Stations are placed at

closer intervals near the coast and islands to accommodate these

features (see Kramer et al., 1972 for further details).

clamping the net line to the towing cable with the 45 kg terminal
weight about 10-15 m below the surface. The net was lowered to
ca . 210 m depth by paying out 300 m of wire over a 6 minute
period (35 m of depth/min.). After fishing at depth for 30
seconds, the net was retrieved at 20 m/min. (14 m depth/min.).
The angle of stray of the towing cable was recorded every 3
seconds and maintained at 45° (+3°) by adjusting the ship speed
and course. After reaching the surface, the net was washed down
and the samples preserved in 5% formalin buffered with sodium
borate. Flowmeter readings were made at the beginning and end of
each tow. Detailed descriptions of gear and methods are given by
Kramer et al . (1972), and Smith and Richardson (1977).

LABORATORY PROCEDURES

Laboratory processing began with the determination of a
displacement volume for each sample (methods described in Staff,
SPFI, 1953 and Kramer et al., 1972). Sorting involved the
removal of ichthyoplankton from the sample and identification and
separation of: eggs and larvae of Pacific sardine and northern
anchovy; larvae of Pacific hake; and eggs of Pacific saury. Some
samples were fractioned into aliquots using a Folsom plankton
splitter (McEwen et al., 1954) prior to sorting. Criteria for
fractioning were: 1) samples taken at a distance greater than
200 nautical miles from shore were not fractioned, 2) samples
taken closer than 200 miles from shore and containing 25 ml of
plankton or less were not fractioned, and 3) samples taken closer
than 200 miles from shore and containing more than 25 ml of
plankton were fractioned to 50% of their original volume (J. R.
Thrailkill, pers. comm.). Aliquot percentages for fractioned
samples from 1975 are listed in Table 1 under the "Percent
Sorted" column; 67.9% of samples collected in 1975 were
fractioned.

A "standard haul factor" (SHF) was calculated for each tow
to make them comparable and allow estimations of areal abundance.
This factor adjusts the number of eggs or larvae in a haul to the
number in 10 m of water strained per meter of depth fished. If
the vertical distribution of the species has been encompassed.
then the adjusted value is equivalent to the number under 10 m
of sea surface. The SHF is calculated for each haul by the
formula :

SHF = 10 D
V

where D = depth of haul = cosine of the average angle
of stray of the towing cable multiplied by
cable length (m)

3
V = total volume of water (m ) strained

during the haul

V = R • a • p

where R = total number of revolutions of the current
meter during the haul

2
a = area (m ) of the mouth of the net

p = length of column of water (m) needed to
produce one revolution of the current

meter.

Tow depth, volume of water strained, and standard haul factor are
listed in Table 1 for each tow taken during 1975. Detailed
descriptions of factors involved in calculating these values are
presented in Ahlstrom (1948), Kramer et al . (1972), and Smith and
Richardson (1977) .

IDENTIFICATION

Identification of ichthyoplankton species beyond those
separated during the sorting process was carried out by a
separate group of specialists. Ontogenetic stages of fishes are
inherently difficult to identify and this is further complicated
by the large number and diversity of species which contribute to
the ichthyoplankton of the California Current region. Most
identifications were accomplished by establishing ontogenetic
series on the basis of morphology, meristics, and pigmentation
and then identifying these series by relating them to known
metamorphic, juvenile, or adult stages with overlapping features
(Powles and Markle, 1984) . A total of 151 taxa was identified
for 1975, with 97 taken to species, 25 to genus, 24 to family,
and 5 to order or suborder. In the decade of the 1970 's some
taxa were identified for the first time. These included larvae
of the bathylagid Bathylagus longirostris , the gonostomatids
Danaphos oculatus and Valencienellus stellatus , the myctophid
Bolinichthys spp. , and the trichiurid Lepidopus xantusi . Larvae
in the families Scopelarchidae and Nomeidae were identified to
genus or species. Five species of rockfish in the Sebastes group
were also identified: S. aurora, S. jordani, S. levis, S.
macdonaldi , and S. paucispinis.

The task of producing a reliable and equitable
ichthyoplankton data base required extensive procedures to
verify, correct, and edit the original identifications. The
primary data source was the original identification sheets (see
Kramer et al . , 1972, for examples); however, a critical resource
used in all phases of this process was the CalCOFI
ichthyoplankton collection in which the samples are archived.
Throughout the course of CalCOFI ichthyoplankton studies, samples
have been identified to the lowest taxon possible. In reviewing
these identifications for the data base, our approach has been
conservative and we have preserved those identifications and
counts which we could confirm, while correcting as many of the
errors as possible. After computer entry, taxonomic errors and

inconsistencies in the data base were corrected and the most
obvious identification errors were corrected. Our current
knowledge of ichthyoplankton techniques coupled with a precise
understanding of the development of identification competency in
the program over the years allowed us to critically judge the
historical records. Identifications were changed to different
taxa, lumped to a higher taxonomic category, or given a more
precise taxonomic name. In some cases, identifications of a
taxon were inconsistent among cruises in a year. These records
were made equitable by lumping to the higher taxonomic category
to avoid biases that could result in quantitative misinter-
pretation.

Next, statistical, seasonal, and geographic outliers were
identified, employing a series of graphic summaries and listings.
Examination of geographic outliers proved to be especially
effective because of our accumulated knowledge of species
distributions. In the course of examining samples for these
outliers, other identification errors were discovered and
eventually all taxa were scrutinized to some extent. Lastly,
certain taxa were reexamined in all samples for the entire
CalCOFI time series. These taxa were selected because of their
commercial, ecological, phylogenetic, or zoogeographic importance
or because taxonomic confusion was at the ordinal level. The
following is a list of the taxa for 1975 which received special
attention, with explanations and caveats intended to aid in
quantitative interpretations:

Anguillif ormes - tentative and sporadic identifications to family
or lower taxon lumped to order.

Sardinops sagax - all specimens south of line 120 checked for
misidentif ication of Opisthonema spp.

Engraulis mordax - some nearshore samples of small E. mordax
may contain other anchovy genera which could not be
differentiated.

Nansenia spp. - all specimens checked and identified as N .

Candida or N. crassa ; all specimens of these species near
their range boundaries checked.

Bathylagus spp. - includes small and/or disintegrated specimens
of Bathylagus or Leuroglossus stilbius .

Stomiiformes - all specimens checked and identified to genus
or species; residuals are small, poorly preserved or
unavailable specimens.

Cyclothone spp. - tentative and sporadic identifications to
species were lumped to genus.

Vinciguerria lucetia - some V. poweriae may remain in these

samples because small larvae of the two species could not be

differentiated; sporadic identification of V. poweriae
began in 1961.

Sternoptychidae - tentative and sporadic identifications of
hatchetf ishes to genus were lumped to family.

Paralepididae - all specimens examined and identified to species;
residuals are small, poorly preserved or unavailable
specimens .

Scopelarchidae - all specimens reidentified to species except
Scopelarchus ; residuals are a small, poorly preserved
specimen and one unavailable specimen.

Scopelarchus spp. - tentative and sporadic identifications to
species lumped to genus.

Lampanyctus spp. - tentative and sporadic identifications to
species lumped to genus.

Lampanyctus regal is - underrepresented because of inability

to differentiate small larvae (<5 mm) from those of other
species of the genus; counts may include other species of
the genus because of difficulty in identifying larvae of
this large and complex genus.

Lampanyctus ritteri - comment for L. regal is applies to this
species .

Diogenichthys atlanticus - all specimens at margins of range
checked.

Diogenichthys laternatus - all specimens at margins of range
checked.

Hygophum spp. - all specimens reidentified to species.

Hygophum atratum - all specimens checked.

Hygophum reinhardtii - all specimens checked.

Symbolophorus calif orniensis - all specimens south of line 120
checked.

Ophidiiformes - this category did not exist originally and

unidentified larvae of this order, including a type referred
to as "Zoarcidae", were originally placed in the "blenny"
category.

Chilara taylori - all specimens checked.

Ophidion scrippsae - all specimens checked.

Trachipteridae - tentative and sporadic identifications to genus
were lumped to family.

Melamphaes spp. - all identifications ascribed to Melamphaidae
were reexamined and assigned to genus (Melamphaes ,
Poromitra) or species (Scopelogadus bispinosus) ; larvae
originally identified as Melamphaes spp. were not reexamined
and this category may contain other melamphaid genera.

Anoplopoma fimbria - specimen checked.

Cottidae - all specimens checked.

Oxylebius pictus - all specimens checked.

Zaniolepis spp. - all specimens checked.

Blennioidei - this is the residual of the completely reexamined
"blenny" category, which also contained various misidenti-
fied ophidiif orms, and is now restricted to members of
northern stichaeioid families and true blennioids (other
than Hypsoblennius spp.) in the southern part of the
pattern) .

Labridae - all specimens originally identified to family were
reexamined and assigned to genus (Halichoeres spp.) or
species (Oxyjulis calif ornica , Semicossyphus pulcher) .

Chromis punctipinnis - records south of about line 120 may
include other pomacentrid taxa.

Howella brodiei - specimen checked; in this report we list H.
brodiei in the family Apogonidae for convenience,
recognizing that its systematic affinities are not resolved.

Carangidae - all specimens checked; tentative and sporadic
identifications to genus or species (except Trachurus
symmetricus and Seriola lalandi) were lumped to family.

Seriola lalandi - all specimens checked.

Gerreidae - tentative and sporadic identifications to genus
lumped to family.

Haemulidae - tentative and sporadic identifications to genus
lumped to family.

Girella nigricans - specimen checked.

Medialuna calif orniensis - all specimens checked.

Caulolatilus princeps - all specimens checked.

Sciaenidae - tentative and sporadic identifications to genus
lumped to family.

Scombridae - all larvae identified to this family or constituent
taxa (except Scomber japonicus) were reexamined and reas-
signed; residuals are small, poorly preserved specimens.

Pleuronectif ormes - all specimens of this category were
reexamined and reidentif ied.

Bothidae - all specimens examined and reassigned; most were
assigned to various paralichthyid genera.

Citharichthys spp. - all larvae identified to species were lumped
to genus except C. stigmaeus ; category includes larvae of
Etropus spp .

Citharichthys stigmaeus - includes larvae larger than ca. 4.5 mm;
smaller larvae are in Citharichthys spp.

Paralichthys calif ornicus - all specimens examined.

Xystreurys liolepis - originally misidentif ied as Paralichthys
calif ornicus ; all specimens reidentif ied.

Glyptocephalus zachirus - all specimens examined.

Hypsopsetta guttulata - some specimens were originally identified
as Pleuronichthys spp.

Lepidopsetta bilineata - specimens originally identified as
Psettichthys melanostictus .

Microstomus pacificus - all specimens examined.

Pleuronichthys spp. - all larvae of this genus and constituent

species were examined and assigned to species; residuals are
unavailable specimens.

Psettichthys melanostictus - all specimens examined.

COMPUTER ENTRY AND EDITING

Each taxon on the original identification sheets was given a
3-digit code based on the list of codes in Haight et al. (1979).
Taxon codes and counts from these sheets were keypunched by
cruise and station, along with pertinent station and tow data and
entered into the VAX 11/780 computer at the University of
California, San Diego, Computing Center. After entries were
completed for an entire year, print-out listings of taxa and
counts on each station were compared with the original data
sheets to eliminate keypunch errors. Next, data in the file were
cross-checked with data on an existing file which contained:
station and tow data; numbers of eggs of sardine, anchovy, and
saury; numbers of larvae of sardine, anchovy, hake, jack
mackerel, and Pacific mackerel; total number of fish eggs; and
total number of fish larvae.

Discrepancies in ichthyoplankton data in these two files
were corrected by inspecting original records from the sorting
laboratory, the original ichthyoplankton identification sheets,
and the samples themselves. Station and tow data discrepancies
between the two files were corrected by reviewing ships' logs and
deck tow sheets, original records from the sorting laboratory,
cruise announcements, publications, header information on the
ichthyoplankton identification sheets, and station plots
generated for each cruise. Eventually all station and tow data
were checked by comparing these sources.

The corrected ichthyoplankton data base was then examined
statistically and outliers were found and checked as above.
Distributional plots were then prepared for each taxon and these
were checked by reviewing the data sources mentioned above and by
examining archived specimens. A listing of each taxon by station
(Table 4) was produced, which became the primary document for
subsequent checks. Misidentif ications found in geographic
outlier checks and other misidentif ications and data problems
discovered in the course of examining archived samples resulted
in several iterations of Table 4. Finally, totals in Table 4
were checked against annual summaries of incidence and abundance
(Tables 2 and 3) . Ecological analyses of the data were conducted
concurrently with editing procedures and provided cross-checks
that allowed correction of errors.

SPECIES SUMMARY

Collections made in 1975 were analyzed separately from those
taken in November and December of 1974 with respect to the pooled
occurrences and counts of larvae (Tables 2A,B; 3A,B). Larvae of
northern anchovy (Engraulis mordax) represented 72% of all fish
larvae taken on CalCOFI cruises during 1975 and numbered almost
10 times as many as Pacific hake, Merluccius productus , the next
most abundant taxon with 7.3% of the total larvae (Table 2b, 3b).
Northern anchovy also ranked first in incidence; M. productus
ranked 8th. The next most abundant was the rockfish genus
Sebastes with 3.9% of the total, followed by the sanddab genus
Citharichthys with 3.0%; they ranked 2nd and 4th respectively in
incidence. The deepsea smelt Leuroglossus stilbius ranked 5th in
abundance (2.5%) and 3rd in occurrence. Two myctophids,
Triphoturus mexicanus and Stenobrachius leucopsarus , ranked 6th
(1.6%) and 7th (1.2%) in number, and 6th and 5th in occurrence.
The final 3 taxa in the top 10 collected in 1975 were the croaker
family Sciaenidae, with 0.9%, the gonostomatid Vinciguerria
lucetia , with 0.7%, and the deepsea smelt Bathylagus ochotensis ,
with 0.6% of total larvae. These 3 taxa ranked 10th, 16th, and
9th in incidence. The appearance of croaker larvae in the top 10
may reflect the increased number of stations occupied on the
shoreward end of each line where these larvae are most abundant.
These 10 top taxa contributed 93.5% to the total number of larvae
collected in 1975; the remaining 6.5% was distributed among 140
taxa plus the disintegrated and unidentified categories. The top

10

10 taxa comprised 4 coastal demersal taxa,
species, and 5 midwater species.

1 coastal pelagic

Six of the 10 most numerous taxa collected in
2A) were also among the top 10 in 1975 - Engraulis
63.4% of the total larvae, Sebastes spp.
spp . (3.7%), Vinciguerria lucetia (2.6%),
Leuroglossus stilbius (2.0%). These six
2nd, 8th, 6th, and 7th in incidence respectively
taxa ranking in the top 10 for 1974 were

1974 (Table

mordax with

(9.7%), Citharichthys

Sciaenidae (2.2%), and

taxa ranked 1st, 3rd,

in 1974. Other

3 myctophids,

Diogenichthys laternatus (1.4%), Protomyctophum crockeri (1.4%),
and Stenobrachius leucopsarus (1.0%); and a rockfish, Sebastes

paucispims
number of

(1.0%). These 10 taxa contributed
larvae collected in the 272 tows
remaining 12.5% was distributed among 78
disintegrated and unidentified categories. Of

87.5% to the total

during 1974; the

taxa plus the

the top 10 taxa, 4

were coastal demersal taxa,
5 were midwater species.

1 was a coastal pelagic species,

and

EXPLANATION OF TABLES

Table 1 - This table lists by cruise the pertinent station and
tow data for 197 5 (including November and December,
1974), the volume of water filtered and standard haul
factor for each tow, the percent of sample sorted, and
the total numbers of fish eggs and larvae. CalCOFI
cruises are designated by four digits; the first two
indicate the year and the second two the month.
Within each cruise the data are listed in order of
increasing line and station number (southerly and
seaward directions) ; the order of station occupancy is
shown on the station charts (Figures 2-10) . Stations
are designated by two groups of digits; the first set
indicates the line and decimal fraction and the second
set indicates the station on the line. Time is listed
as Pacific Standard Time at the start of each tow in
24-hour designation. Methods for determining tow
depth, volume of water strained, standard haul factor,
and percent sorted were described in the methods
section. The values for total fish eggs and larvae
represent raw counts (unadjusted for percent sorted or
standard haul factor). Ship codes are as follows: JD,
David Starr Jordan; AX, Alexander Agassi z .

Table 2A- This table lists pooled occurrences of all larval fish
taxa taken during November and December of 1974 in
ranked order.

Table 2B- This table lists pooled occurrences of all larval
taxa taken during 1975 in ranked order.

fish

Table 3A- This table lists pooled counts of all larval fish taxa
taken during November and December of 1974 in ranked

11

order. Numbers are adjusted for percent sorted and
standard haul factors.

Table 3B- This table lists pooled counts of all larval fish taxa
during 1975 in ranked order. Numbers are adjusted for
percent sorted and standard haul factors.

Table 4 - This table gives numbers of fish larvae for each taxon
in 1974 and 1975, listed by station and calendar month
in which the tow was taken. Counts are adjusted for
percent of sample sorted and standard haul factor.
Average values are given for stations occupied more
than once during a month. See Table 1 for station and
tow data and Table 6 for listing of stations with
multiple occupancies during a month. Multiple
occupancies occurred when a station was occupied more
than once during a calendar month; in some cases,
multiple occupancies resulted from separate cruises.
The orders are listed in "phylogenetic" seguence
modified from Nelson (1984) . Subtaxa within each order
are listed alphabetically. Page numbers for each taxon
are given in the index at the end of the report.

Table 5 - This table is a summary of pooled occurrences of all
larval fish taxa taken on CalCOFI surveys from 1972 to
1981. Taxa are listed in the same order as in Table

4.

Table 6 - List of stations with multiple occupancies in one month
during the 1975 survey.

ACKNOWLEDGMENTS

Elbert Ahlstrom, John Butler, Susan D'Vincent, Connie Fay,
Barbara MacCall, Geoff Moser, Elaine Sandknop, and Betsy Stevens
originally identified larvae from CalCOFI cruises of 1975.
Ronald Whyte coded each larval fish taxon or type and Rita Ford
entered them into the computer. Debby Snow efficiently assisted
in all aspects of data editing and retrieval. Cindy Meyer and
James Ryan provided programming assistance. Dorothy Roll
designed the CalCOFI data acguisition system and provided data
processing support. Ken Raymond, Roy Allen, and Henry Orr helped
with graphics and production of the report. Lorraine Prescott
prepared the manuscript for printing. Paul Smith determined
statistical outliers, provided assistance during geographical
outlier checks and offered helpful suggestions throughout the
project. Izadore Barrett, Director of the Southwest Fisheries
Center provided support critical to the completion of the
project. James Thrailkill planned CalCOFI surveys and supervised
cruises, data handling, and plankton sorting from 1949 to 1986
and is largely responsible for the high quality of these
operations. Without the vision and direction of Elbert Ahlstrom
and Elton Sette and the dedicated efforts of the many people who
collected, processed, and analyzed the samples, this data base

12

would not exist. During the final stages of preparing this
report, Reuben Lasker succumbed to cancer. As Chief of the
Coastal Fisheries Resources Division, SWFC, his encouragement and
support for all of us involved in the sea surveys, sample
processing, and data base and report preparation were unwavering.
We dedicate this work to his memory.

13

LITERATURE CITED

Ahlstrom, E. H. 1948. A record of pilchard eggs and larvae
collected during surveys made in 1939 to 1941. U.S. Fish
Wildl. Serv. SSRF 54, 82 p.

Ahlstrom, E. H., H. G. Moser, and E. M. Sandknop. 1978.
Distributional atlas of fish larvae in the California
Current region: rockfishes, Sebastes spp. , 1950 through
1975. CalCOFI Atlas No. 26:xxi + 178 p.

Ambrose, D. A., R. L. Charter, H. G. Moser, and C. R. Santos
Methot. 1987a. Ichthyoplankton and station data for
California Cooperative Oceanic Fisheries Investigations
survey cruises in 1951. U.S. Dep. Commer., NOAA Tech.
Memo., NMFS, SWFC, No. 79, 196 p.

Ambrose, D. A., R. L. Charter, H. G. Moser, and C. R. Santos
Methot. 1987b. Ichthyoplankton and station data for
California Cooperative Oceanic Fisheries Investigations
survey cruises in 1955. U.S. Dep. Commer., NOAA Tech.
Memo., NMFS, SWFC, No. 83, 18 5 p.

Ambrose, D. A., R. L. Charter, H. G. Moser, and C. R. Santos
Methot. 1987c. Ichthyoplankton and station data for
California Cooperative Oceanic Fisheries Investigations
survey cruises in 1960. U.S. Dep. Commer., NOAA Tech.
Memo., NMFS, SWFC, No. 88, 2 53 p.

Ambrose, D. A., R. L. Charter, H. G. Moser, and B. S. Earhart.

1988a. Ichthyoplankton and station data for California

Cooperative Oceanic Fisheries Investigations survey cruises

in 1963. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 94, 209 p.

Ambrose, D. A., R. L. Charter, H. G. Moser, and B. S. Earhart.
1988b. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1967. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 98, 103 p.

Haight, C. A., H. G. Moser, and P. E. Smith. 1979. Data entry
programs: CalCOFI. II. Fish eggs and larvae identification
sheet. National Marine Fisheries Service, Southwest
Fisheries Center, La Jolla, Admin. Rept . No. LJ-79-25.

Hewitt, R. 1980. Distributional atlas of fish larvae in the
California Current region: northern anchovy, Engraulis
mordax Girard, 1966 through 1979. CalCOFI Atlas No. 28:
xi + 101 p.

14

Kramer, D. , M. Kalin, E. G. Stevens, J. R. Thrailkill, and J. R.
Zweifel. 1972. Collecting and processing data on fish eggs
and larvae in the California Current Region. NOAA Tech.
Rep. NMFS Circ. 370, 38 p.

McEwen, G. F., M. W. Johnson, and T. R. Folsom. 1954. A
statistical analysis of the performance of the Folsom
Plankton Sample Splitter, based on test observations. Arch.
Meteor. Geophys . Bioklim. Ser. A, 7:502-527.

Nelson, J. S. 1984. Fishes of the world. John Wiley and Sons,
N.Y. , 523 p.

Powles, H. and D. F. Markle. 1984. Identification of larvae, p.
31-33. In: Ontogeny and systematics of fishes. H. G.
Moser, W. J. Richards, D. M. Cohen, M. P. Fahay, A. W.
Kendall, Jr., and S. L. Richardson (eds.). Spec. Publ .
No. 1. Amer. Soc. Ichthyol . Herpetol., 760 p.

Sandknop, E. M. , R. L. Charter, H. G. Moser, and J. D. Ryan.
1987a. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1952. U.S. Dep . Commer. , NOAA Tech. Memo., NMFS, SWFC,
No. 80, 207 p.

Sandknop, E. M. , R. L. Charter, H. G. Moser, and J. D. Ryan.
1987b. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1958. U.S. Dep. Commer. NOAA Tech. Memo., NMFS, SWFC,
No. 86, 248 p.

Sandknop, E. M. , R. L. Charter, H. G. Moser, C. A. Meyer, and A.
E. Hays. 1988a. Ichthyoplankton and station data for
California Cooperative Oceanic Fisheries Investigations
survey cruises in 1961. U.S. Dep. Commer., NOAA Tech.
Memo., NMFS, SWFC, No. 92, 167 p.

Sandknop, E. M., R. L. Charter, H. G. Moser, C. A. Meyer, and A.
E. Hays. 1988b. Ichthyoplankton and station data for
California Cooperative Oceanic Fisheries Investigations
survey cruises in 1964. U.S. Dep. Commer., NOAA Tech.
Memo., NMFS, SWFC, No. 95, 222 p.

Sandknop, E. M. , R. L. Charter, H. G. Moser, C. A. Meyer, and A.
E. Hays. 1988c. Ichthyoplankton and station data for
California Cooperative Oceanic Fisheries Investigations
survey cruises in 1968. U.S. Dep. Commer., NOAA Tech.
Memo., NMFS, SWFC, No. 99, 112 p.

Smith, P. E. 1974. Distribution of zooplankton volumes in the
California Current region, 1969. CalCOFI Atlas No. 20:xv-
xviii + 118-125.

15

Smith, P. E. and S. L. Richardson. 1977. Standard techniques
for pelagic fish egg and larva surveys. FAO Fish. Tech.
Pap. No. 175, 100 p.

Staff, South Pacific Fishery Investigations. 1953. Zooplankton
volumes off the Pacific Coast, 1952. U.S. Fish Wildl. Serv.
SSRF 100, 41 p.

Stevens, E. G. , R. L. Charter, H. G. Moser, and M. S. Busby.
1987a. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1953. U.S. Dep. Commer. , NOAA Tech. Memo., NMFS , SWFC,
No. 81, 186 p.

Stevens, E. G., R. L. Charter, H. G. Moser, and M. S. Busby.
1987b. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1956. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 84, 189 p.

Stevens, E. G., R. L. Charter, H. G. Moser, and M. S. Busby.
1987c. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1959. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 87, 273 p.

Stevens, E. G., R. L. Charter, H. G. Moser, and L. R. Zins,
1988a. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1965. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 96, 220 p.

Stevens, E. G., R. L. Charter, H. G. Moser, and L. R. Zins.
1988b. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1969. U.S. Dep. Commer., NOAA Tech. memo., NMFS, SWFC,
No. 100, 265 p.

Sumida, B. Y., R. T>. Charter, H. G. Moser, and D. L. Snow.
1987a. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1954. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 82, 207 p.

Sumida, B. Y., R. L. Charter, H. G. Moser, and D. L. Snow.
1987b. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 19 57. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 85, 225 p.

Sumida, B. Y., R. L. Charter, H. G. Moser, and D. L. Snow.
1988a. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises

16

in 1962. U.S. Dep . Commer. , NOAA Tech. Memo, NMFS , SWFC,
No. 93, 179 p.

Sumida, B. Y., R. L. Charter, H. G. Moser, and D. L. Snow.

1988b. Ichthyoplankton and station data for California

Cooperative Oceanic Fisheries Investigations survey cruises

in 1966. U.S. Dep. Commer., N07AA Tech. Memo., NMFS, SWFC,
No. 97, 287 p.

Sumida, B. Y., R. L. Charter, H. G. Moser, and D. L. Snow.
1988c. Ichthyoplankton and station data for California
Cooperative Oceanic Fisheries Investigations survey cruises
in 1972. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC,
No. 109, 219 p.

University of California, Scripps Institution of Oceanography.
1984a. Data report: physical and chemical data, CalCOFI
Cruises 7412, 7501, 7503. SIO Ref. 84-10.

University of California, Scripps Institution of Oceanography.
1984b. Data report: physical and chemical data, CalCOFI
Cruises 7505, 7507, 7510, 7512. SIO Ref. 84-13.

17

7412

7501

7510

\

7503

7505

V

7511

7512

Figure 1. Composite arrangement of diagrammatic charts showing
areas sampled on each CalCOFI cruise during 1975.

18

t 1 r

20°

CRUISE 7412

DAVID STARR JORDAN

20"

125°

120°

110°

Figure 2

Station pattern for CalCOFI Cruise 7412 showing the
track for the David Starr Jordan. Stations with
plankton tows are indicated by a dot; circles
designate hydrographic stations; diamonds signify STD
recordings. Figures 2-10 modified from charts in
Univ. of Calif., SIO (1984a, 1984b) to include only
those stations listed in Table 1 of this report.

19

125°

-i 1 r

115° 110°

1 1 1 1 1

40°

35°

30°

25°

20°

T T

CRUISE 7412

ALEXANDER AGASSIZ

40°

J I I L

i : ,

J L

20"

120°

H5°

Figure 3. Station pattern for CalCOFI Cruise 7412 - Alexander
Agassi z. Symbols as in Figure 2.

20

CRUISE 7501

40°

35°

30°

DAVID STARR JORDAN

25°

20°

20"

'25°

20°

Figure 4. Station pattern for CalCOFI Cruise 7501 - David
Starr Jordan. Symbols as in Figure 2.

21

T 1 r

15° 110°

1 1 1 1

40'

35'

30°

25'

20

t

CRUISE 7501

ALEXANDER AGASSIZ

J L

J I I L

I I i

J I I I L

30"

20"

125"

120°

115°

H0°

Figure 5. Station pattern for CalCOFI Cruise 7501 - Alexander
Agassi z. Symbols as in Figure 2.

22

40'

35'

30°

t 1 r

i 1 1 r

CRUISE 7503

ALEXANDER AGASSIZ

25"

20°

J L

DAVID STARR JORDAN

J 1 I L

J_J I L

J I L

20'

125°

120°

115°

110°

Figure 6. Station pattern for CalCOFI Cruise 7503. Symbols as
in Figure 2.

23

1 1 I

125°

120°

15°

CRUISE 7505

40°

35°

30°

DAVID STARR JORDAN

110°

—

25°

40°

20°

J L

J L

2C

125°

120°

115°

u0°

Figure 7. Station pattern for CalCOFI Cruise 7505
in Figure 2.

Symbols as

24

40'

35'

30°

25'

t r

i 1 r

t 1 r

CRUISE 7507

20'

- DAVID STARR JORDAN *-§,

ALEXANDER AGASSIZ

20°

125°

120°

115°

110°

Figure 8. Station pattern for CalCOFI Cruise 7507. Symbols as
in Figure 2. "P" and "N" denote phytoplankton and
neuston tows, respectively; these data were not
included in this report.

25

40'

if

30°

t r

CRUISES 7510 AND 751 I

25'

DAVID STARR JORDAN

CRUISE 751 I

CRUISE 7510

20'

J L

I l i

-i i

20*

125°

120"

1 15*

1

i;G°

Figure 9. Station pattern for CalCOFI Cruises 7510 and 7511
Symbols as in Figure 8.

26

"5° no-

"1 1 1 1 1

CRUISE 7512

40°

35"

30°

25°

DAVID STARR JORDAN

20°

125°

20°

H0°

Figure 10. Station pattern for CalCOFI Cruise 7512. Net tow
stations indicated by dots; see Univ. of Calif.,
SIO, 1984b for hydrographic stations.

27

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CAPE

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* • • . . *1

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Figure 11. The basic station plan for CalCOFI cruises from 1950
to the present.

28

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TABLE 2A. Pooled occurrences of fish larvae taken during CalCOFI
cruises in 1974.

Rank Taxon Occurrences

1 Engraulis mordax 155

2 Citharichthys spp. 96

3 Sebastes spp. 94

4 Sternoptychidae 63

5 Protomyctophum crockeri 6 2

6 Sciaenidae 58

7 Leuroglossus stilbius 52

8 Vinciguerria lucetia 48

9 Triphoturus mexicanus 38

10 Lampanyctus spp. 3 5

11 Citharichthys stigmaeus 33

12 Cyclothone spp. 3

13 Diogenichthys laternatus 29
13 Stenobrachius leucopsarus 29

15 Disintegrated fish larva 27

16 Tarletonbeania crenularis 26
16 Gobiidae 26

18 Paralichthys calif ornicus 2 5

19 Diogenichthys atlanticus 22

20 Unidentified fish larva 21

21 Idiacanthus antrostomus 18

22 Lestidiops ringens 16
2 2 Merluccius productus 16

2 4 Bathylagus wesethi 15

25 Symbolophorus calif orniensis 14

26 Bathylagus ochotensis 13

27 Myctophidae 12

28 Lampanyctus ritteri 11
28 Sardinops sagax 11
30 Chauliodus macouni 10
30 Sebastes paucispinis 10
30 Gonostomatidae 10

3 3 Gonichthys tenuiculus 9
3 3 Melamphaes spp. 9

3 3 Stomias atriventer 9
33 Clinidae 9
37 Hippoglossina stomata 8
37 Nansenia crassa 8
37 Symphurus spp. 8
37 Microstoma microstoma 8
37 Tetragonurus cuvieri 8

4 2 Synodus spp. 7
42 Trachipteridae 7
42 Rosenblattichthys volucris 7
45 Ophidion scrippsae 6
45 Peprilus simillimus 6
45 Hypsoblennius spp. 6
45 Myctophum nitidulum 6

67

TABLE 2A. ( cont . )

Rank Taxon Occurrences

45 Danaphos oculatus 6

45 Diogenichthys spp. 6

45 Argentina sialis 6

45 Icichthys lockingtoni 6

45 Hygophum atratum 6

45 Parophrys vetulus 6

55 Hypsopsetta guttulata 5

55 Ceratoscopelus townsendi 5

55 Paralepididae 5

55 Cottidae 5

55 Chiasmodontidae 5

55 Bathylagus milleri 5

61 Scopelogadus bispinosus 4

61 Xystreurys liolepis 4

63 Atherinidae 3

6 3 Syngnathus spp. 3

6 3 Macroramphosus gracilis 3

6 3 Scorpaenichthys marmoratus 3

67 Anguilliformes 2

67 Lampadena urophaos 2

67 Zaniolepis spp. 2

67 Pleuronichthys ritteri 2

71 Ceratioidei 1

71 Sebastolobus spp. 1

71 Pleuronichthys decurrens 1

71 Macropinna microstoma 1

71 Scopelosaurus spp. 1

71 Lepidopus xantusi 1

71 Cyclopteridae 1

71 Ichthyococcus spp. 1

71 Pleuronichthys spp. 1

71 Agonidae 1

71 Lampanyctus regalis 1

71 Blennioidei 1

71 Coryphaena hippurus 1

71 Microstomus pacificus

71 Sefcastes jordani 1

71 Stomiif ormes 1

71 Bathylagus spp. 1

71 Bathylagus pacificus 1

71 Pleuronichthys verticalis 1

71 Cololabis saira 1

68

TABLE 2B. Pooled occurrences of fish larvae taken during CalCOFI
cruises in 1975.

Rank Taxon Occurrences

1 Engraulis mordax 8 42

2 Sebastes spp. 5 60

3 Leuroglossus stilbius 363

4 Citharichthys spp. 357

5 Stenobrachius leucopsarus 351

6 Triphoturus mexicanus 342

7 Protomyctophum crockeri 29 9

8 Merluccius productus 27 9

9 Bathylagus ochotensis 273

10 Sciaenidae 260

11 Sternoptychidae 218

12 Tarletonbeania crenularis 215

13 Disintegrated fish larva 196

14 Unidentified fish larva 183

15 Cyclothone spp. 16 5

16 Vinciguerria lucetia 164

17 Bathylagus wesethi 156

18 Lampanyctus spp. 151

19 Lampanyctus ritteri 149

20 Diogenichthys atlanticus 141

21 Citharichthys stigmaeus 133
2 2 Melamphaes spp. 130
23 Gobiidae 121

2 4 Symbolophorus calif orniensis 120

25 Trachurus symmetricus 119

26 Diogenichthys laternatus 114

27 Paralichthys calif ornicus 106

28 Pleuronichthys verticalis 100

29 Uypsoblennius spp. 82

30 Myctophidae 80

31 Chauliodus macouni 7 8

3 2 Sebastes paucispinis 7 3
3 3 Diaphus spp. 7
34 Clinidae 67

3 5 Ceratoscopelus townsendi 66
36 Stomias atriventer 59
36 Argentina sialis 59

38 Serranidae 55

39 Peprilus simillimus 54

40 Sardinops sagax 51

41 Parophrys vetulus 5

4 2 Danaphos oculatus 4 9
4 3 Icichthys lockingtoni 46
44 Cottidae 44
4 5 Sebastes jordani 4 2
46 Synodus spp. 41
46 Bathylagus spp. 41
4 8 Microstoma microstoma 40

69

TABLE 2B. (cont . )

Rank Taxon Occurrences

49 Bathylagus pacificus 39

49 Lestidiops ringens 39

51 Hippoglossina stomata 36

5 2 Pleuronichthys ritteri 3 3

53 Idiacanthus antrostomus 30

54 Lampanyctus regalis 29

55 Symphurus spp. 2 6
5 5 Halichoeres spp. 26

5 5 Nansenia Candida 26
58 Rosenblattichthys volucris 23
58 Zaniolepis spp. 2 3
58 Oxyjulis calif ornica 2 3
58 Sebastolobus spp. 23
62 Myctophum nitidulum 22

6 2 Chromis punctipinnis 2 2

64 Sebastes macdonaldi 21

65 Lyopsetta exilis 20
6 6 Scopelarchus spp. 19
67 Trachipteridae 18
6 7 Ophidion scrippsae 18

6 7 Poromitra spp. 18

7 Sebastes levis 17
70 Paralepididae 17
7 Nansenia crassa 17
70 Chilara taylori 17
7 4 Bygophum atratum 16
7 5 Tetragonurus cuvieri 15
75 Ophidiif ormes 15
7 5 Scorpaenichthys marmoratus 15
7 5 Etrumeus acuminatus 15
7 5 Diogenichthys spp. 15

80 Gonichthys tenuiculus 14

81 Bathylagus milleri 13
81 Cyclopteridae 13
81 Sebastes aurora 13
81 Notolychnus valdiviae 13
85 Prionotus spp. 12
85 Xystreurys liolepis 12
85 Gonostomat idae 12
88 Chiasmodontidae 11
88 Ceratioidei 11
88 Agonidae 11
88 Scorpaena spp. 11
9 2 Scopelosaurus spp. 10
92 Lepidopus xantusi 10
92 Carangidae 10
92 Gobiesocidae 10
96 Sphyraena argentea 9
96 Microstomus pacificus 9

70

TABLE 2B. (cont. )

Rank Taxon Occurrences

96 Hygophum reinhardtii 9

9 9 Scomber japonicus 8

99 Anguillif ormes 8

9 9 Semicossyphus pulcher 8

99 Hypsopsetta guttulata 8

99 Syngnathus spp. 8

99 Haemulidae 8

9 9 Ichthyococcus spp. 8

106 Electrona rissoi 7

106 Cololabis saira 7

106 Atherinidae 7

109 Notoscopelus resplendens 6

110 Brosmophycis marginata 5
110 Seriola lalandi 5
110 Lampadena urophaos 5
110 Notolepis risso 5
110 Scopelogadus bispinosus 5
110 Gerreidae 5
116 Oxylebius pictus 4
116 Glyptocephalus zachirus 4
116 Blennioidei 4
116 Coryphaena hippurus 4
120 Benthalbella dentata 3
120 Benthosema pterota 3
120 Loweina rara 3
120 Sarda chiliensis 3
120 Medialuna calif orniens is 3
120 Pleuronichthys decurrens 3
120 Macrouridae 3
120 Hypsypops rubicundus 3
120 Lepidopsetta bilineata 3
120 Brama spp. 3
120 Pleuronichtbys coenosus 3
131 Scopelarchidae 2
131 Psettichthys melanostictus 2
131 Caulolatilus princeps 2
131 Aristostomias scintillans 2
131 Scope larchoides nicholsi 2
136 Valenciennellus stellatus 1
136 Pleuronichthys spp. 1
136 Anoplopoma fimbria 1
136 Scombridae 1
136 Icosteus aenigmaticus 1
136 Macropinna microstoma 1
136 Photonectes spp. 1
136 Diplophos taenia 1
136 Polynemidae 1
136 Girella nigricans 1
136 Platichthys stellatus 1

71

TABLE 2B. ( cont . )

Rank Taxon Occurrences

136 Bowel la brodiei 1

136 Exocoetidae 1

136 Opisthonema spp. 1

136 Ophiodon elongatus 1

136 Hexagrammidae 1

136 Stomiiformes 1

72

TABLE 3A. Pooled numbers of fish larvae taken during CalCOFI

cruises in 1974. Counts are adjusted for percent of
sample sorted and standard haul factor (see text).

Rank Taxon Count

1 Engraulis mordax 39 366

2 Sebastes spp. 6042

3 Citharichthys spp. 2 306

4 Vinciguerria lucetia 1604

5 Sciaenidae 1350

6 Leuroglossus stilbius 1246

7 Diogenichthys laternatus 893

8 Protomyctophwn crockeri 8 88

9 Stenobrachius leucopsarus 64 3

10 Sebastes paucispinis 628

11 Sternoptychidae 613

12 Citharichthys stigmaeus 552

13 Lampanyctus spp. 490

14 Tarletonbeania crenularis 487

15 Triphoturus mexicanus 371

16 Sardinops sagax 2 89

17 Bathylagus ochotensis 281

18 Cyclothone spp. 2 80

19 Diogenichthys atlanticus 261

2 Merluccius productus 20 9

21 Paralichthys calif ornicus 204

22 Disintegrated fish larva 194

23 Gobiidae 177

24 Unidentified fish larva 153

25 Idiacanthus antrostomus 147

26 Bathylagus wesethi 141

27 Symbolophorus calif orniensis 124

28 Gonichthys tenuiculus 100

29 Chauliodus macouni 97

30 Stomias atriventer 91

31 Lestidiops ringens 89

32 Myctophidae 84

3 3 Diogenichthys spp. 7 6
3 3 Lampanyctus ritteri 7 6
3 5 Icichthys lockingtoni 7 5
3 5 Melamphaes spp. 7 5
37 Trachipteridae 70

3 8 Synodus spp. 6 9

39 Myctophum nitidulum 68

40 Argentina sialis 66

41 Symphurus spp. 6 5

4 2 Microstoma microstoma 60

43 Bathylagus milleri 58

44 Rosenblattichthys volucris 57

45 Macroramphosus gracilis 52

46 Peprilus simillimus 51
46 Parophrys vetulus 51
46 Clinidae 51

73

TABLE 3A. (cont

Rank Taxon Count

49 Bygophum atratum 48

50 Gonostomatidae 45

51 Nansenia crassa 40

52 Paralepididae 39

5 2 Tetragonurus cuvieri 39

54 Xystreurys liolepis 37

55 Ceratoscopelus townsendi 3 6

56 Chiasmodontidae 32
56 Cottidae 32

58 Lampadena urophaos 3

59 Hypsoblennius spp. 2 9
59 Danaphos oculatus 2 9
61 Hypsopsetta guttulata 22

6 2 Hippoglossina stomata 21
6 3 Ophidion scrippsae 17

64 Agonidae 13
6 5 Scopelogadus bispinosus 12

65 Bathylagus pacificus 12
65 Blennioidei 12
6 5 Microstomus pacificus 12

6 5 Lampanyctus regalis 12
65 Atherinidae 12
71 Pleuronichthys decurrens 11
71 Cololabis saira 11

7 3 Zaniolepis spp. 9
7 4 Scorpaenichthys marmoratus 8
7 4 Sebastes jordani 8
7 6 Coryphaena hippurus 6
7 6 Syngnathus spp. 6
78 Anguillif ormes 5
78 Pleuronichthys ritteri 5
80 Macropinna microstoma 3
80 Sebastolobus spp. 3
80 Bathylagus spp. 3
80 Lepidopus xantusi 3
80 Cyclopteridae 3
80 Ceratioidei 3
80 Ichthyococcus spp. 3
80 Stomiiformes 3
80 Scopelosaurus spp. 3
89 Pleuronichthys spp. 2
89 Pleuronichthys verticalis 2

Total 62101

74

TABLE 3B. Pooled numbers of fish larvae taken during CalCOFI

cruises in 1975. Counts are adjusted for percent of
sample sorted and standard haul factor (see text).

Rank Taxon Count

1 Engraulis mordax 838883

2 Merluccius productus 84347

3 Sebastes spp. 45007

4 Citharichthys spp. 34806

5 Leuroglossus stilbius 28735

6 Triphoturus mexicanus 18081

7 Stenobrachius leucopsarus 14 507

8 Sciaenidae 10537

9 Vinciguerria lucetia 847 3

10 Bathylagus ochotensis 7171

11 Trachurus symmetricus 487 5

12 Protomyctophum crockeri 3911

13 Tarletonbeania crenularis 3880

14 Citharichthys stigmaeus 3828

15 Bathylagus wesethi 3470

16 Diogenichthys laternatus 3082

17 Sebastes jordani 2907

18 Sternoptychidae 2627

19 Cyclothone spp. 2441
2 Sardinops sagax 2 360

21 Lampanyctus ritteri 2174

22 Disintegrated fish larva 1998

23 Unidentified fish larva 1946

24 Diogenichthys atlanticus 1658

2 5 Sebastes paucispinis 1599

26 Lampanyctus spp. 1369

27 Gobiidae 1327

28 Paralichthys calif ornicus 1151

29 Diaphus spp. 1132

3 Melamphaes spp. 10 50

31 Symbolophorus calif orniensis 1048

32 Hypsoblennius spp. 1032

33 Pleuronichthys verticalis 961

3 4 Peprilus simillimus 898

35 Clinidae 872

36 Serranidae 827

37 Parophrys vetulus 798

38 Argentina sialis 714

39 Myctophidae 685

40 Chromis punctipinnis 683

41 Sarda chiliensis 670

42 Cottidae 620

4 3 Symphurus spp. 596

44 Ceratoscopelus townsendi 571

45 Stomias atriventer 556

46 Chauliodus macouni 553

47 Ealichoeres spp. 44 3

75

TABLE 3B. (cont. )

Rank Taxon Count

48 Bathylagus pacificus 439

49 Synodus spp. 42 8

50 Bathylagus spp. 409

51 Carangidae 400

52 Danaphos oculatus 364

53 Icichthys lockingtoni 358
5 4 Sebastes macdonaldi 332

5 5 Sebastes levis 3 28

56 Etrumeus acuminatus 270

57 Lampanyctus regalis 264

58 Hippoglossina stomata 254

59 Microstoma microstoma 2 47

60 Zaniolepis spp. 240

61 Lyopsetta exilis 235

6 2 Sebastolobus spp. 2 29

6 2 Lestidiops ringens 229

64 Pleuronichthys ritteri 210

65 Nansenia Candida 202

66 Ophidiif ormes 193

67 Idiacanthus antrostomus 180

68 Oxyjulis calif ornica 157

69 Chilara taylori 150

70 Bathylagus milleri 148

71 Ophidion scrippsae 147

7 2 Scorpaenichthys marmoratus 146
7 3 Scorpaena spp. 138
7 4 Nansenia crassa 137
75 Paralepididae 130
7 6 Sebastes aurora 109
77 Xystreurys liolepis 106
7 8 Prionotus spp. 103

79 Trachipteridae 100

80 Poromitra spp. 99

81 Anguilliformes 98

82 Tetragonurus cuvieri 97

83 Diogenichthys spp. 95

84 Hygophum atratum 94
84 Myctophum nitidulum 94

86 Gonichthys tenuiculus 90

87 Agonidae 86

88 Haemulidae 84

89 Notolychnus valdiviae 82

90 Scopelarchus spp. 81

91 Lepidopus xantusi 8

92 Rosenblattichthys volucris 78
9 3 Semicossyphus pulcher

94 Gonostomat idae 74

95 Atherinidae 73
95 Seriola lalandi 73

76

TABLE 3B. (cont . )

Rank Taxon Count

97 Microstomas pacificus 71

98 Gobiesocidae 66

99 Cyclopteridae 63

100 Scomber japonicus 6 2

101 Bypsopsetta guttulata 61
101 Sphyraena argentea 61

103 Brosmophycis marginata 57

104 Benthosema pterota 5 4
10 5 Scopelosaurus spp. 5 3

106 Chiasmodontidae 51

107 Cololabis saira 49

108 Ceratioidei 48

109 Ichthyococcus spp. 46
109 Bygophum reinhardtii 46
111 Medialuna californiensis 4 5
111 Blennioidei 45

113 Caulolatilus princeps 43

114 Syngnathus spp. 41

115 Notoscopelus resplendens 40

116 Glyptocephalus zachirus 39

117 Oxylebius pictus 38

118 Macrouridae 37

119 Pleuronichthys decurrens 35

120 Bypsypops rubicundus 3 4

121 Gerreidae 31

122 Psettichthys melanostictus 29

123 Benthalbella dentata 28

124 Scombridae 26

125 Lepidopsetta bilineata 25

126 Lampadena urophaos 2 2
126 Coryphaena hippurus 2 2

128 Electrona rissoi 2

129 Brama spp. 18

130 Notolepis risso 17
130 Scopelogadus bispinosus 17
132 Valenciennellus stellatus 14
132 Scopelarchidae 14

134 Girella nigricans 13

135 Exocoetidae 12
135 Macropinna microstoma 12
137 Icosteus aenigmaticus 11
137 Stomi if ormes 11
137 Polynemidae 11
137 Platicbthys stellatus 11

141 Hexagrammidae 10

142 Loweina rara 9

143 Pleuronichthys coenosus 8

144 Scopelarchoides nicholsi 1

145 Aristostomias scintillans 6

77

TABLE 3B. (cont. )

Rank

Taxon

Count

146
147
147
147
150
150
150

Pleuronichthys spp.
Ophiodon elongatus
Diplophos taenia
Photonectes spp.
Anoplopoma fimbria
Opisthonema spp.
Howella brodiei

5
3
3
3
2
2
2

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1162305

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TABLE 6. List of stations which were occupied twice in
one month during 1975.

Station

Mon

93.0

60.0

11

63.0

50.0

2

63.0

52.0

2

63.0

55.0

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70.0

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60.0

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65.0

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70.0

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73.0

50.0

2

(1974)

224

INDEX

This index lists taxa included in Table 4 with their page numbers.

Page

Anguilliformes 79

Clupei formes

Clupeidae

Etrumeus acuminatus 79

Opistonema spp 7 9

Sardinops sagax 7 9

Engraulidae

Engraulis mordax 81

Salmoni formes

Argentinidae

Argentina sialis 86

Microstoma microstoma 88

Nansenia Candida 88

Nansenia crassa 89

Bathylagidae

Bathylagus spp 89

Bathylagus milleri 90

Bathylagus ochotensis 91

Bathylagus pacificus 94

Bathylagus wesethi 9 5

Leuroglossus stilbius 97

Opisthoproctidae

Macropinna microstoma 101

Stomiiformes 102

Gonostomatidae 102

Cyclothone spp 102

Danaphos oculatus 104

Diplophos taenia 105

Ichthyococcus spp 105

Valenciennellus stellatus 106

Vinciguerria lucetia 106

Sternoptychidae 108

Stomiatoidea

Chaul iodont idae

Chauliodus macouni Ill

Idiacanthidae

Idiacanthus antrostomus 113

Malacosteidae

Aristostomias scintillans 113

Mel anostomi idae

Photonecte spp 114

Stomiidae

Stomias atriventer 114

Myctophi formes
Alepisauroidei

225

Page

Paralepididae 115

Lestidi ops ringens 115

Notolepis risso 117

Chloropthalmoidei

Notosudidae

Scopel osaurus spp 117

Scopelarchidae 117

Benthalbella dentata 117

Rosenblattichthys volucris 117

Scopel archoides nicholsi 118

Scopelarchus spp 118

Myctophoidei

Myctophidae 119

Lampanyctinae

Ceratoscopel us townsendi 12

Diaphus spp 121

Lampadena urophaos 12 3

Lampanyctus spp 123

Lampanyctus regalis 12 5

Lampanyctus ritteri 12 6

Notolychnus valdiviae 128

Notoscopel us resplendens 129

Stenobrachius leucopsarus 129

Triphoturus mexicanus 13 3

Myctophinae

Benthosema pterota 136

Diogenichthys spp 137

Diogenichthys atlanticus 137

Diogenichthys laternatus 139

Electrona rissoi 14

Gonichthys tenuiculus 141

Hygophum atratum 141

Hygophum reinhardtii 142

Loweina rara 142

Myctophum nitidulum 142

Protomyctophum crockeri 14 3

Symbolophorus calif orniensis 146

Tarletonbeania crenularis 148

Synodontoidei

Synodontidae

Synodus spp 150

Gadiformes

Merlucciidae

Merluccius productus 151

Macrouridae 155

Ophidiif ormes 155

Bythitidae

Brosmophycis marginata 156

Ophidiidae

Chilara taylori 156

Ophidion scrippsae 157

226

Page

Lophiiformes

Ceratioidei 157

Gobiesoci formes

Gobiesocidae 157

Bel on i formes

Exocoetidae 158

Scomberesocidae

Cololabis saira 158

Atherini formes

Atherinidae 158

Lampri formes

Trachipteridae 158

Beryci formes

Melamphaidae

Melamphaes spp 159

Poromi tra spp 161

Scopelogadus bispinosus 162

Syngnathi formes

Macroramphos idae

Macroramphosus gracilis 162

Syngnathidae

Syngnathus spp 162

Scorpaeni formes
Cottoidei

Agonidae 162

Anoplopomatidae

Anoplopoma fimbria 163

Cottidae 163

Scorpaeni chthys marmoratus 164

Cyclopteridae 164

Hexagrammidae 165

Ophiodon elongatus 165

Oxylebius pictus 165

Zani ol epi s spp 165

Scorpaenoidei

Scorpaenidae

Scorpaena spp 166

Sebastes spp 166

Sebastes aurora 170

Sebastes jordani 171

Sebastes levis 172

Sebastes macdonaldi 172

Sebastes paucispinis 173

Sebastolobus spp 174

Triglidae

Prionotus spp 175

Perciformes

Blennioidei 175

Blenniidae

Hypsobl enni us spp 175

Clinidae 177

Gobioidei

227

Page

Gobiidae 178

Icosteoidei

Icosteidae

Icosteus aenigmaticus 180

Labroidei

Labridae

Halichoeres spp 180

Oxyjulis calif ornica 181

Semicossyphus pulcher 182

Pomacentridae

Chromis punctipinnis 182

Hypsypops rubicundus 182

Percoidei

Apogonidae

Howell a brodiei 183

Bramidae

Brama spp 183

Carangidae 183

Seri ola lal andi 183

Trachurus symmetricus 183

Coryphaenidae

Coryphaena hippurus 185

Gerreidae 185

Haemulidae 186

Kyphosidae

Girella nigricans 186

Medial una calif orniensis 186

Malacanthidae

Caulolatilus princeps 186

Sciaenidae 186

Serranidae 189

Polynemidae 191

Scombroidei

Scombridae 191

Sarda chiliensis 191

Scomber j aponi cus 191

Trichiuridae

Lepidopus xantusi 191

Sphyraenoidei

Sphyraenidae

Sphyraena argentea 192

Stromateoidei

Centrolophidae

Icichthys lockingtoni 192

Stromateidae

Peprilus simillimus 193

Tetragonuridae

Tetragonurus cuvieri 194

Trachinoidei

Chiasmodontidae 194

Pleuronecti formes
Pleuronectoidei

Paral ichthy idae

228

Page

Citharichthys spp 195

Citharichthys stigmaeus 200

Hippoglossina stomata 202

Paralichthys calif ornicus 203

Xystreurys liolepis 205

Pleuronectidae

Glyptocephalus zachirus 205

Hypsopsetta guttulata 2 05

Lepidopsetta bilineata 206

Lyopsetta exilis 206

Microstomias pacificus 2 06

Parophrys vetul us 2 07

Platichthys stellatus 2 08

Pleuronichthys spp 2 08

Pleuronichthys coenosus 2 08

Pleuronichthys decurrens 208

Pleuronichthys ritteri 2 09

Pleuronichthys verticalis 209

Psettichthys melanost ictus 211

Soleoidei

Cynoglossidae

Symph urus spp 211

Disintegrated fish larva 212

Unidentified fish larva 215

229

*U.S. GPO 1988/584-471/34010/7-21-88/175

RECENT TECHNICAL MEMORANDUMS

Copies of this and other NOAA Technical Memorandums are available from the National Technical
Information Service, 5285 Port Royal Road, Springfield, VA 22167. Paper copies vary in price.
Microfiche copies cost $4.50. Recent issues of NOAA Technical Memorandums from the NMFS
Southwest Fisheries Center are listed below:

NOAA-TM-NMFS-SWFC- 100 Ichthyoplankton and station data for California Cooperative

Oceanic Fisheries Investigations survey cruises in 1969.
E.G. STEVENS, R.L CHARTER, H.G. MOSER, and LR. ZINS
(January 1988)

101 Hawaiian monk seal population monitoring, pup captive
maintenance program, and incidental observations of the green
turtle at Kure Atoll, 1985.

M.L REDDY and C.A. GRIFFITH
(January 1988)

102 Hydrographic observations in the northwestern Weddell Sea
marginal ice zone during March 1986.

D.M. HUSBY and R.D. MUENCH
(January 1988)

103 Deep-sea shrimp trapping for Heterocarpus laevigatus in the
Hawaiian Archipelago by a commercial fishing vessel.

D.T. TAGAMI and S. BARROWS
(March 1988)

104 Report of ecosystem studies conducted during the 1986 eastern
tropical Pacific dolphin survey on the research vessel McArthur.
V.G. THAYER, B.G. McDONALD, J.M. ELLINGSON, C.W. OLIVER,
D.W. BEHRINGER and S.B. REILLY

(March 1988)

105 Report of ecosystem studies conducted during the 1986 eastern
tropical Pacific dolphin survey on the research vessel David Starr
Jordan.

V.G. THAYER, R.L PITMAN, K.A. RITTMASTER, G.G. THOMAS,
D.W. BEHRINGER and S.B. REILLY
(March 1988)

1 06 An economic analysis of lobster fishing vessels performance in the
Northwestern Hawaiian Islands.

R.P. CLARK and S.G. POOLEY
(April 1988)

107 The Hawaiian monk seal and green turtle on Pearl and Hermes
Reef, 1986.

R.G. FORSYTH, D.J. ALCORN, T. GERRODETTE and W.G.

GILMARTIN

(April 1988)

108 A review of California entangling net fisheries, 1981-1986.
S.F. HERRICK, JR. and D. HANAN

(June 1988)

109 Ichthyoplankton and station data for California Cooperative
Oceanic Fisheries Investigations survey cruises in 1972.
B.Y. SUMIDA, R.L. CHARTER, H.G. MOSER and D.L SNOW
(June 1988)1